The Caytoniales (Figs. 1-2) are an extinct order of seed plants known from fossils collected throughout the Mesozoic Era, specifically in the late Triassic to Maastrichtian period, around 250 to 70 million years ago. They are regarded as seed ferns because they are seed-bearing plants with fern-like leaves. Although at one time considered angiosperms because of their berry-like cupules, that hypothesis was later disproven. Nevertheless, some authorities consider them likely ancestors of angiosperms, whereas others consider angiosperms more likely derived from Glossopteridales. The origin of angiosperms remains an intriguing puzzle (see Evolutionary history of plants).